Publications

The following publications represent research conducted by or in collaboration with the Field Research Centre (FRC):

Casaer, J., Wevers, J., Bollen, M., Desmet, P., Govaert, S., Milotic, T., Indestege, S., Kuypers, W., Tomsin, I., & Beenaerts, N. (2026). NPHK1 - Systematic random camera trap observations in National Park Hoge Kempen (Belgium). https://doi.org/10.15468/suawqp

Gordillo-Pérez, M. J., Reyes-Tur, B., Chen, Z., Sigwart, J., Greve, C., Heleven, M., Smeets, K., & Beenaerts, N. (2026). Genome-wide identification and evolutionary characterization of predicted HSP70 proteins in the Cuban painted snail Polymita picta. Frontiers in Ecology and Evolution, 14. https://doi.org/10.3389/fevo.2026.1933916

Dornelas, M., Antão, L. H., Bates, A. E., Brambilla, V., Chase, J. M., Chow, C. F. Y., Fontrodona-Eslava, A., Magurran, A. E., Martins, I. S., Moyes, F., Sagouis, A., Adu-Acheampong, S., Acquah-Lamptey, D., Adam, D., Ajani, P. A., Albaina, A., Almaraz, P., An, J., Anderson, R. S., … Zvuloni, A. (2025). BioTIME 2.0: Expanding and improving a database of biodiversity time series. Global Ecology and Biogeography, 34(5). https://doi.org/10.1111/geb.70003

Gordillo-Pérez, M. J., Beenaerts, N., Sánchez, D. L., Smeets, K., Arias-Sosa, Y. C., & Reyes-Tur, B. (2025). Shell colour luminance of Cuban painted snails, Polymita picta and Polymita muscarum (Gastropoda: Cepolidae). PLOS ONE, 20(1), e0314008. https://doi.org/10.1371/journal.pone.0314008

Indestege, S., Bollen, M., Casaer, J., & Beenaerts, N. (2025). Do long-term camera trap sampling designs matter to estimate diel activity? European Journal of Wildlife Research, 71(6). https://doi.org/10.1007/s10344-025-02013-3

Kuypers, W., Bollen, M., Casaer, J., & Beenaerts, N. (2025). Trapped by trails: How different types of recreational trails influence seasonal space use of wildlife in a densely visited national park. Science of The Total Environment, 996, 180091. https://doi.org/10.1016/j.scitotenv.2025.180091

Lathouwers, M., Beenaerts, N., Evens, R., Artois, T., Conway, G., Henderson, I., Shewring, M., Cross, T., Ulenaers, E., & Dendoncker, N. (2025). Impact of future climate change and land-use change on habitat suitability for a long-distance avian migrant under diverse socioeconomic-emission scenarios. Global Ecology and Conservation, 62, e03777. https://doi.org/10.1016/j.gecco.2025.e03777

Nouwen, O., Rineau, F., Kohout, P., Baldrian, P., Eisenhauer, N., Beenaerts, N., Thijs, S., Vangronsveld, J., & Soudzilovskaia, N. A. (2025). Towards understanding the impact of mycorrhizal fungal environments on the functioning of terrestrial ecosystems. FEMS Microbiology Ecology, 101(8). https://doi.org/10.1093/femsec/fiaf062

Pärtel, M., Tamme, R., Carmona, C. P., Riibak, K., Moora, M., Bennett, J. A., Chiarucci, A., Chytrý, M., de Bello, F., Eriksson, O., Harrison, S., Lewis, R. J., Moles, A. T., Öpik, M., Price, J. N., Amputu, V., Askarizadeh, D., Atashgahi, Z., Aubin, I., … Zobel, M. (2025). Global impoverishment of natural vegetation revealed by dark diversity. Nature, 641(8064), 917–924. https://doi.org/10.1038/s41586-025-08814-5

Tomsin, I., Duthie, A. B., Bunnefeld, N., Leirs, H., Casaer, J., & Beenaerts, N. (2025). Evaluating management scenarios for the european hamster ( Cricetus cricetus) using quantitative models. Ecology and Evolution, 15(10). https://doi.org/10.1002/ece3.72353

Bollen, M., Casaer, J., Neyens, T., & Beenaerts, N. (2024). When and where? Day-night alterations in wild boar space use captured by a generalized additive mixed model. PeerJ, 12, e17390. https://doi.org/10.7717/peerj.17390

Burton, A. C., Beirne, C., Gaynor, K. M., Sun, C., Granados, A., Allen, M. L., Alston, J. M., Alvarenga, G. C., Calderón, F. S. Á., Amir, Z., Anhalt-Depies, C., Appel, C., Arroyo-Arce, S., Balme, G., Bar-Massada, A., Barcelos, D., Barr, E., Barthelmess, E. L., Baruzzi, C., … Kays, R. (2024). Mammal responses to global changes in human activity vary by trophic group and landscape. Nature Ecology &Amp; Evolution, 8(5), 924–935. https://doi.org/10.1038/s41559-024-02363-2

Cruz Portorreal, Y., Beenaerts, N., Koedam, N., Reyes Dominguez, O. J., Milanes, C. B., Dahdouh-Guebas, F., & Pérez Montero, O. (2024). Perception of mangrove social–ecological system governance in southeastern cuba. Water, 16(17), 2495. https://doi.org/10.3390/w16172495

Martijn, B., Thomas, N., Natalie, B., & Jim, C. (2024). Impacts of zoning and landscape structure on the relative abundance of wild boar assessed through a Bayesian N-mixture model. Science of The Total Environment, 911, 168546. https://doi.org/10.1016/j.scitotenv.2023.168546

Bollen, M., Palencia, P., Vicente, J., Acevedo, P., Del Río, L., Neyens, T., Beenaerts, N., & Casaer, J. (2023). Assessing trends in population size of three unmarked species: A comparison of a multi-species N-mixture model and random encounter models. Ecology and Evolution, 13(10). https://doi.org/10.1002/ece3.10595

Jacobs, Jeffrey, Beenaerts, N., & Artois, T. (2023). Green roofs and pollinators, useful green spots for some wild bee species (Hymenoptera: Anthophila), but not so much for hoverflies (Diptera: Syrphidae). Scientific Reports, 13(1). https://doi.org/10.1038/s41598-023-28698-7

Jacobs, J., Verheyde, F., Dekoninck, W., De Smedt, P., Van Dijck, T., Beenaerts, N., & Artois, T. (2023). The importance of green roofs in urban macroinvertebrate biodiversity: A case study in Antwerp, Belgium. Research Square.

Lathouwers, M., Dendoncker, N., Artois, T., Beenaerts, N., Conway, G., Henderson, I., Shewring, M., Cross, T., Ulenaers, E., & Evens, R. (2023). Multi-scale habitat selection throughout the annual cycle of a long-distance avian migrant. Ecological Indicators, 156, 111099. https://doi.org/10.1016/j.ecolind.2023.111099

Martijn, B., Jim, C., Natalie, B., & Thomas, N. (2023). Simulation-based assessment of the performance of hierarchical abundance estimators for camera trap surveys of unmarked species. Scientific Reports, 13(1). https://doi.org/10.1038/s41598-023-43184-w

Neyens, T., Petrof, O., Faes, C., Vandenrijt, W., Ulenaers, P., Artois, T., Beenaerts, N., & Evens, R. (2023). Winter agri-environment schemes and local landscape composition influence the distribution of wintering farmland birds. Global Ecology and Conservation, e02533. https://doi.org/10.1016/j.gecco.2023.e02533

Rineau, F., Reyns, W., Carpentier, C., Van Der Plas, F., Bardgett, R., Beenaerts, N., & De Laender, F. (2023). Fungal pairwise interactions shift from positive to negative under warming stress. https://doi.org/10.5194/egusphere-egu23-1282

Jacobs, J., Berg, M., Beenaerts, N., & Artois, T. (2022). Biodiversity of collembola on green roofs: A case study of three cities in Belgium. Ecological Engineering, 177, 106572. https://doi.org/10.1016/j.ecoleng.2022.106572

Lathouwers, M., Artois, T., Dendoncker, N., Beenaerts, N., Conway, G., Henderson, I., Kowalczyk, C., Davaasuren, B., Bayrgur, S., Shewring, M., Cross, T., Ulenaers, E., Liechti, F., & Evens, R. (2022). Rush or relax: Migration tactics of a nocturnal insectivore in response to ecological barriers. Scientific Reports, 12(1), 4964. https://doi.org/10.1038/s41598-022-09106-y

Lathouwers, M., Nussbaumer, R., Liechti, F., Davaasuren, B., Artois, T., Beenaerts, N., Dendoncker, N., Ulenaers, E., & Evens, R. (2022). Migration routes and timing of European Nightjars (Caprimulgus europaeus) breeding in eastern Mongolia. Journal of Ornithology, 163(4), 881–890. https://doi.org/10.1007/s10336-022-02000-4

Bollen, M., Neyens, T., Fajgenblat, M., De Waele, V., Licoppe, A., Manet, B., Casaer, J., & Beenaerts, N. (2021). Managing african swine fever: Assessing the potential of camera traps in monitoring wild boar occupancy trends in infected and non-infected zones, using spatio-temporal statistical models. Frontiers in Veterinary Science, 8. https://doi.org/10.3389/fvets.2021.726117

Roy, J., Rineau, F., De Boeck, H. J., Nijs, I., Pütz, T., Abiven, S., Arnone, J. A., Barton, C. V. M., Beenaerts, N., Brüggemann, N., Dainese, M., Domisch, T., Eisenhauer, N., Garré, S., Gebler, A., Ghirardo, A., Jasoni, R. L., Kowalchuk, G., Landais, D., … Milcu, A. (2021). Ecotrons: Powerful and versatile ecosystem analysers for ecology, agronomy and environmental science. Global Change Biology, 27(7), 1387–1407. https://doi.org/10.1111/gcb.15471

Wevers, J., Beenaerts, N., Casaer, J., Zimmermann, F., Artois, T., & Fattebert, J. (2021). Modelling species distribution from camera trap by-catch using a scale-optimized occupancy approach. Remote Sensing in Ecology and Conservation, 7(3), 534–549. https://doi.org/10.1002/rse2.207

Boix, Y. F., Dubois, A. E. F., Hendrix, S., Luna, L. M. G., Beenaerts, N., Manrique, C. E. M., Victório, C. P., & Cuypers, A. (2020). Assessment of the antioxidative potential of rosmarinus officinalis L. (Lamiaceae) irrigated with static magnetic field-treated water. Brazilian Archives of Biology and Technology, 63. https://doi.org/10.1590/1678-4324-2020190142

Evens, R., Conway, G., Franklin, K., Henderson, I., Stockdale, J., Beenaerts, N., Smeets, K., Neyens, T., Ulenaers, E., & Artois, T. (2020). DNA diet profiles with high-resolution animal tracking data reveal levels of prey selection relative to habitat choice in a crepuscular insectivorous bird. Ecology and Evolution, 10(23), 13044–13056. https://doi.org/10.1002/ece3.6893

Llauradó Maury, G., Méndez Rodríguez, D., Hendrix, S., Escalona Arranz, J. C., Fung Boix, Y., Pacheco, A. O., García Díaz, J., Morris-Quevedo, H. J., Ferrer Dubois, A., Aleman, E. I., Beenaerts, N., Méndez-Santos, I. E., Orberá Ratón, T., Cos, P., & Cuypers, A. (2020). Antioxidants in plants: A valorization potential emphasizing the need for the conservation of plant biodiversity in cuba. Antioxidants, 9(11), 1048. https://doi.org/10.3390/antiox9111048

Pilotto, F., Kühn, I., Adrian, R., Alber, R., Alignier, A., Andrews, C., Bäck, J., Barbaro, L., Beaumont, D., Beenaerts, N., Benham, S., Boukal, D. S., Bretagnolle, V., Camatti, E., Canullo, R., Cardoso, P. G., Ens, B. J., Everaert, G., Evtimova, V., … Haase, P. (2020). Meta-analysis of multidecadal biodiversity trends in Europe. Nature Communications, 11(1). https://doi.org/10.1038/s41467-020-17171-y

Vanderkelen, Inne, Zscheischler, J., Gudmundsson, L., Keuler, K., Rineau, F., Beenaerts, N., Vangronsveld, J., Vicca, S., & Thiery, W. (2020). A novel method for assessing climate change impacts in ecotron experiments. International Journal of Biometeorology, 64(10), 1709–1727. https://doi.org/10.1007/s00484-020-01951-8

Vanderkelen, I., Zscheischler, J., Gudmundsson, L., Keuler, K., Rineau, F., Beenaerts, N., Vangronsveld, J., Vicca, S., & Thiery, W. (2020). Correction to: A novel method for assessing climate change impacts in ecotron experiments (International Journal of Biometeorology, (2020), 64, 10, (1709-1727), 10.1007/S00484-020-01951-8). International Journal of Biometeorology, 64(10), 1729.

Wevers, J., Fattebert, J., Casaer, J., Artois, T., & Beenaerts, N. (2020). Trading fear for food in the Anthropocene: How ungulates cope with human disturbance in a multi-use, suburban ecosystem. Science of The Total Environment, 741, 140369. https://doi.org/10.1016/j.scitotenv.2020.140369

Boix, Y. F., Dubois, A. F., Alemán, E. I., Victório, C. P., do Carmo de Oliveira Arruda, R., Cuypers, A., Beenaerts, N., Aguilera, J. G., & Zuffo, A. M. (2019). Static magnetic treatment of irrigation water on differents plants cultures improving development. In Estudos interdisciplinares nas ciências exatas e da terra e engenharias (pp. 76–84). Atena Editora. https://doi.org/10.22533/at.ed.2181911097

Neyens, T., Diggle, P. J., Faes, C., Beenaerts, N., Artois, T., & Giorgi, E. (2019). Mapping species richness using opportunistic samples: A case study on ground-floor bryophyte species richness in the Belgian province of Limburg. Scientific Reports, 9(1). https://doi.org/10.1038/s41598-019-55593-x

Norevik, G., Åkesson, S., Artois, T., Beenaerts, N., Conway, G., Cresswell, B., Evens, R., Henderson, I., Jiguet, F., & Hedenström, A. (2019). Wind-associated detours promote seasonal migratory connectivity in a flapping flying long-distance avian migrant. Journal of Animal Ecology, 89(2), 635–646. https://doi.org/10.1111/1365-2656.13112

Reyns, W., Rineau, F., Spaak, J. W., Franken, O., Berg, M. P., Van Der Plas, F., Bardgett, R. D., Beenaerts, N., & De Laender, F. (2019). Food web uncertainties influence predictions of climate change effects on soil carbon sequestration in heathlands. Microbial Ecology, 79(3), 686–693. https://doi.org/10.1007/s00248-019-01444-1

Rineau, F., Malina, R., Beenaerts, N., Arnauts, N., Bardgett, R. D., Berg, M. P., Boerema, A., Bruckers, L., Clerinx, J., Davin, E. L., De Boeck, H. J., De Dobbelaer, T., Dondini, M., De Laender, F., Ellers, J., Franken, O., Gilbert, L., Gudmundsson, L., Janssens, I. A., … Vangronsveld, J. (2019). Towards more predictive and interdisciplinary climate change ecosystem experiments. Nature Climate Change, 9(11), 809–816. https://doi.org/10.1038/s41558-019-0609-3

Vanderkelen, I., Zschleischler, J., Gudmundsson, L., Keuler, K., Rineau, F., Beenaerts, N., Vangronsveld, J., & Thiery, W. (2019). A new approach for assessing climate change impacts in ecotron experiments. https://doi.org/10.5194/bg-2019-267

Beenaerts, N., Evens, R., Beenaerts, N., Neyens, T., Witters, N., Smeets, K., & Artois, T. (2018). Proximity of breeding and foraging areas affects foraging effort of a crepuscular, insectivorous bird. Scientific Reports (Nature Publishing Group). https://doi.org/10.1038/s41598-018-21321-0

Beenaerts, N., Rineau, F., Janssens, I., Steegen, P., Roland, M., Arriga, N., Vanderkelen, I., Ceulemans, R., Thiery, W., Vangronsveld, J., Gudmunsson, L., Zscheischler, J., Davin, E., & Seneviratne, S. I. (2018). From climate change to weather conditions: Linking ecotrons to an ecosystem measurement tower to better simulate the effect of climate change on ecosystem functioning.

Boix, Y. F., Alemán, E. I., Torres, J. M., Chávez, E. R., Arruda, R. do C. de O., Hendrix, S., Beenaerts, N., Victório, C. P., Luna, L. G., Manrique, C. M., & Cuypers, A. (2018). Magnetically treated water on phytochemical compounds of Rosmarinus officinalis L. International Journal of Environment, Agriculture and Biotechnology, 3(1), 297–303. https://doi.org/10.22161/ijeab/3.1.37

Evens, R., Beenaerts, N., Witters, N., & Artois, T. (2018). First records of the chewing louse Mulcticola hypoleucus (Denny, 1842) on the Eurasian nightjar Caprimulgus europaeus Linnaeus, 1758 in the Benelux. Belgian Journal of Zoology, 148(1). https://doi.org/10.26496/bjz.2018.17

Beenaerts, N., Evens, R., Beenaerts, N., Witters, N., & Artois, T. (2017). Study on the foraging behaviour of the European nightjar Caprimulgus europaeus reveals the need for a change in conservation strategy in Belgium. Journal of Avian Biology. https://doi.org/10.1111/jav.00996

Evens, R., Beenaerts, N., Witters, N., & Artois, T. (2017). Repeated migration of a juvenile european nightjar caprimulgus europaeus. Journal of Ornithology, 158(3), 881–886. https://doi.org/10.1007/s10336-017-1459-2

Beenaerts, N., Jeanloz, S., Lizin, S., Beenaerts, N., Brouwer, R., Van Passel, S., & Witters, N. (2016). Towards a more structured selection process for attributes and levels in choice experiments: A study in a Belgian protected area. Ecosystem Services. https://doi.org/10.1016/j.ecoser.2016.01.006

Artois, T., Schockaert, E., Beenaerts, N., & Reygel, P. (2013). Trigonostomum vanmecheleni sp. Nov., a new species of Trigonostomidae (Rhabdocoela: Dalytyphloplanida) from the channels of Venice (Italy), with a discussion on theT. Lilliei species group. Italian Journal of Zoology, 80(1), 46–51. https://doi.org/10.1080/11250003.2012.754058

Beenaerts, N., Hensen, W., Cliquet, A., Draye, A. M., Van Passel, S., Natalie, Mertens, K., & Vanheusden, B. (2013). Juridische studie over de bescherming van, en betaling voor, ecosysteemdiensten.

Beenaerts, N., Pethiyagoda, R., Ng, P. K. L., Yeo, D. C. J., Bex, G. J., Bahir, M. M., & Artois, T. (2010). Phylogenetic diversity of Sri Lankan freshwater crabs and its implications for conservation. Molecular Ecology, 19(1), 183–196. https://doi.org/10.1111/j.1365-294x.2009.04439.x

Beenaerts, N. (2009). Molecular phylogenetics and biogeography of the Old World true freshwater crabs, with emphasis on the fauna of Sri Lanka.

Beenaerts, N., & Berghe, E. V. (2007). Comparative study of three transect methods to assess coral cover, richness and diversity. Western Indian Ocean Journal of Marine Science, 4(1). https://doi.org/10.4314/wiojms.v4i1.28471

Beenaerts, N. (2005). Historical biogeography of parathelphusidae on the indian subcontinent. Beenaerts, N., bahir, M., ng, P. K. L., pethiyagoda, R. & Bossuyt, F., 2005,. 12th Benelux Congress of Zoology, Wageningen, the Netherlands - Oral Presentation.

Helgen, K. M., Groves, C. P., Bossuyt, F., Meegaskumbura, M., Beenaerts, N., Gower, D. J., Pethiyacoda, R., Roelants, K., Mannaert, A., Wilkinson, M., Schneider, C. J., Bahir, M. M., Manamendra-Arachchi, K., Ng, P. K. L., Oommen, O. V., & Milinkovitch, M. C. (2005). Biodiversity in sri lanka and the western ghats [1] (multiple letters). Science, 308(5719), 199.

Bossuyt, F., Meegaskumbura, M., Beenaerts, N., Gower, D. J., Pethiyagoda, R., Roelants, K., Mannaert, A., Wilkinson, M., Bahir, M. M., Manamendra-Arachchi, K., Ng, P. K. L., Schneider, C. J., Oommen, O. V., & Milinkovitch, M. C. (2004). Local endemism within the western ghats-sri lanka biodiversity hotspot. Science, 306(5695), 479–481. https://doi.org/10.1126/science.1100167

Meester, L. D., & Beenaerts, N. (1993). Heritable variation in carotenoid content in Daphnia magna. Limnology and Oceanography, 38(6), 1193–1199. https://doi.org/10.4319/lo.1993.38.6.1193